WPF: How can I stretch the middle child in a DockPanel?

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You need to set DockPanel.Dock attached property for your elements and leave TextBox as the last element:

<RadioButton HorizontalAlignment="Stretch"
             HorizontalContentAlignment="Stretch">
    <DockPanel LastChildFill="True">
        <TextBlock DockPanel.Dock="Left"
                   VerticalAlignment="Center"
                   Text="in location:" />
        <Button DockPanel.Dock="Right"
                Margin="10,0,0,0"
                Padding="3,0"
                Content="..." />
        <TextBox Margin="10,0,0,0">
            Path string
        </TextBox>
    </DockPanel>
</RadioButton>

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